I have both the of the 527 Carbines (223 & 7.62) as well as quite a few other 527 rifles. None have given me any primer ignition problems. Some of these get used during Montana's winter as coyote rifles. They see cold weather. The striker/firing pin assemblies also get run dry, no lube. There lubed on the cocking cams and that's it. When I began hunting in northern Wisconsin the Remington 740/760 pump and semi autos were pretty popular. The hunters who used these rifles ran them dry if they wanted the rifle to be reliable. As something of a general rule WD-40 isn't considered a good firearms lubricant. It has one use and that's water displacement from metal hence the WD formula 40 name.
Agreed...
If I understand it, this is the problem. OP's rifle works fine in temps above sub-zero. Below that it shows light primer strikes and rounds that do not fire.
If I am wrong about that please correct me.
Then, when temps fall below that, the rifle has on again, off again, ignition of both milsurp and civilian ammo, with neither one being more prevalent than the other.
So, something is causing the rifle to not ignite the primer of the ammo after temps get really low.
There are TWO possible explanations. One- the rifle has something gumming up the works and causing light strikes at low temps, and Two- the low temps are causing the chemical components of the primers not to ignite as intended.
If the rifle is getting light strikes in warmer weather, then the second possibility is the likely culprit.
Given that the rifle works fine in weather above sub-zero temps, it is highly unlikely that the problem is with the rifle itself. It's either with the affect of sub-zero temps and the ammunition, or something that is gumming the works up in such cold.
So, the only firearm related answer that I see as a possibility, would be a weak firing pin spring. In which case a specialty spring wouldn't necessarily be the answer, just a different spring.
Thats just my two cents. FWIW